Jeffrey D. Muehlbauer Theodore A. Kennedy 20250605 spreadsheet Environmental Data Initiative Environmental Data Initiative Toskey, E.K., T. Matsuoka, P.M. Kiffney, S.M. Naman, M.C. Liermann, J. Bailey, D.A. Beauchamp, J.R. Bellmore, C.A. Binckley, S. Blumenshine, T.J. Caldwell, M.P. Carey, S. Claeson, C. Clawson, R. Cox, K.J. Cromwell, R.J. Danehy, M.J. Dodrill, J.J. Duda, J.B. Fellman, A.Z. Grimm, T.B. Hallbert, C.E. Jordan, E.R. Keeley, B.P. Kennedy, T.A. Kennedy, A.E. Kohler, E.D. Lowery, K. Macneale, K.D. Martens, S.A. Morley, J.D. Muehlbauer, J.R. Neuswanger, A. O'Dowd, S. Pneh, M. Power, S.J. Rhoades, J. Rosenfeld, G. Rossi, B.L. Sanderson, K.K. Shelby, T. Starky-Owens, J.N. Thompson, C. Volk, J.S. Williamshen, and R.W. Wisseman. 2025. DRIFteRS (DRift Invertebrates For salmonids in River Systems): A dataset of drift invertebrate densities in streams and rivers across western North America, 1997–2024 ver 1. Environmental Data Initiative. https://doi.org/10.6073/pasta/234b23f08b48458e7d3910045ebacd9f. https://doi.org/10.6073/pasta/234b23f08b48458e7d3910045ebacd9f These data describe aquatic macroinvertebrate drift downstream of tailwaters below large dams throughout the Colorado River Basin, collected in spring, summer, and fall 2015. Study sites include the tailwaters downstream of Fontenelle Dam (Wyoming) and Flaming Gorge Dam (Utah) on the Green River, Crystal Dam (Colorado) on the Gunnison River, Navajo Dam on the San Juan River (New Mexico), and Glen Canyon, Hoover, Davis, and Parker Dams on the Colorado River (Arizona, Nevada, California). The data also describe seasonal drift in Bright Angel Creek, a tributary of the Colorado River in Grand Canyon, Arizona, from 2015-2017. Data were collected to facilitate comparison of drift concentrations and aquatic biodiversity across tailwaters within the Colorado River Basin and to understand food availability for fishes in Bright Angel Creek. 20150505 20170119 ground condition Complete None planned Range of drift sampling locations -114.7434 -107.6195 42.0261 34.1834 ISO 19115 Topic Category biota USGS Thesaurus macroinvertebrates freshwater ecosystems river systems inland fishery resources USGS Metadata Identifier USGS:79c92c09-6870-44c6-8b85-1a01d2793146 None Colorado River Grand Canyon Green River San Juan River Gunnison River Bright Angel Creek None Diptera Chironomidae Simuliidae Potamopyrgus antipodarum Gammarus lacustris Ephemeroptera Baetidae Trichoptera Hydropsychidae Hydroptilidae Dreissena bugensis Kingdom Animalia animals Subkingdom Bilateria triploblasts Infrakingdom Protostomia Superphylum Ecdysozoa Phylum Arthropoda arthropods Subphylum Hexapoda hexapods Class Insecta insects Subclass Pterygota winged insects Infraclass Neoptera modern, wing-folding insects Superorder Holometabola Order Diptera gnats mosquitoes true flies hoverflies TSN: 118831 Suborder Nematocera long-horned flies Infraorder Culicomorpha Family Chironomidae midges TSN: 127917 Family Simuliidae TSN: 126640 Order Trichoptera caddisflies TSN: 115095 Superfamily Hydropsychoidea Family Hydropsychidae net-spinning caddisflies TSN: 115398 Superfamily Hydroptiloidea Family Hydroptilidae micro-caddisflies TSN: 115629 Infraclass Palaeoptera ancient winged insects Order Ephemeroptera mayflies TSN: 100502 Suborder Pisciforma Family Baetidae TSN: 100755 Subphylum Crustacea crustaceans Superclass Altocrustacea Class Malacostraca Subclass Eumalacostraca Superorder Peracarida Order Amphipoda amphipods Suborder Senticaudata Infraorder Gammarida gammarid amphipods Superfamily Gammaroidea Family Gammaridae Genus Gammarus Species Gammarus lacustris TSN: 93789 Superphylum Spiralia Phylum Mollusca molluscs mollusks Class Gastropoda gastropods slugs snails Subclass Orthogastropoda Infraclass Caenogastropoda Order Littorinida Suborder Rissoidina Superfamily Truncatelloidea Family Tateidae Genus Potamopyrgus Species Potamopyrgus antipodarum New Zealand mud snail TSN: 205006 Class Bivalvia bivalves clams Subclass Autobranchia Infraclass Heteroconchia Order Myida Superfamily Dreissenoidea Family Dreissenidae Genus Dreissena Species Dreissena bugensis quagga mussel TSN: 567514 None. Please see 'Distribution Info' for details. None. Users are advised to read the dataset's metadata thoroughly to understand appropriate use and data limitations. Theodore A. Kennedy USGS Grand Canyon Monitoring and Research Center Research Ecologist mailing and physical

2255 N Gemini Dr

Flagstaff Arizona 86001 USA 928-556-7374 tknnedy@usgs.gov Data were collected in the field under standard protocol, following published methods for QA/QC that involved ensuring minimal net clogging and verifying flow within the next against an external reference. Lab identification and enumeration of invertebrates was carried out by trained technicians with verification by expert taxonomists. Baxter, C.V. T.A. Kennedy, S.W. Miller, J.D. Muehlbauer, and L.A. Smock. 2017. Macroinvertebrate drift, adult insect emergence, and oviposition. pp. 435-456 In F. R. Hauer and G. A. Lamberti, editors. Methods in stream ecology. Academic Press, San Diego. 3rd Edition. Muehlbauer, J. D., Kennedy, T. A., Copp, A. J., and T.A. Sabol. 2017. Deleterious effects of net clogging on the quantification of stream drift. Canadian Journal of Fisheries and Aquatic Sciences, 74(7), 1041-1048. Values fall within expected ranges for environmental and biological data, based on personal experience and visual verification of ranges. Data represent only macroinvertebrate drift and associated environmental conditions collected at the time of drift sampling, within a specific range of dates and locations consistent with project design. Collection of macroinvertebrates was carried out by placing a 250 um mesh net in the water and allowing macroinverterbrates to drift into it. Flow was sampled concurrently, both within and outside the net. Drift sampling procedures followed standard published methods, cited here: Baxter, C.V. T.A. Kennedy, S.W. Miller, J.D. Muehlbauer, and L.A. Smock. 2017. Macroinvertebrate drift, adult insect emergence, and oviposition. pp. 435-456 In F. R. Hauer and G. A. Lamberti, editors. Methods in stream ecology. Academic Press, San Diego. 3rd Edition. Muehlbauer, J. D., Kennedy, T. A., Copp, A. J., and T.A. Sabol. 2017. Deleterious effects of net clogging on the quantification of stream drift. Canadian Journal of Fisheries and Aquatic Sciences, 74(7), 1041-1048. 20170119 Macroinvertebrate enumeration and identification was carried out in a lab setting by trained technicians, using dissecting microscopes at magnification power up to 40x. Sample identifications were verified in the lab by expert taxonomists. Samples were counted their entirety, with no subsampling. Data were entered into a SQL-based database, with automated error checking to ensure data met within previously established ranges for QA/QC. 20190101 ColoradoRiverDriftData.csv Comma Separated Value (CSV) file containing data. Producer Defined SampleID Sample ID Producer Defined Unique name of sample; formatted as ProjectName_Stream_Reach_Date_Replicate. ProjectName Project name, named for location and earliest and latest years sampled. Producer Defined Colorado River Basin Tailwaters 2015 Data from dam tailwater collectionsites. Producer defined Bright Angel Creek 2016-2017 Data from tributary stream collection site. Producer defined Basin Basin name; largest drainage basin that the stream sampled is a tributary to. Producer Defined Colorado River Sites within the Colorado River Basin. Producer defined Stream Stream name. Producer Defined San Juan River The San Juan River downstream of Navajo Dam. Producer defined Green River The Green River downstream of Fontenelle or Flaming Gorge Dams. Producer defined Colorado River The Colorado River downstream of Glen Canyon, Hoover, Davis, and Parker Dams. Producer defined Gunnison River The Gunnison River downstream of Crystal Dam. Producer defined Bright Angel Creek Bright Angel Creek in the Grand Canyon. Producer defined Reach Reach name or number; site name to distinguish between different locations on the same stream or river. Producer Defined SJNavajo San Juan River downstream of Navajo Dam. Producer defined GRFGD Green River downstream of Flaming Gorge Dam Producer defined GRFont Green River downstream of Fontenelle Dam. Producer defined CRLeesFerry Colorado River downstream of Glen Canyon Dam. Producer defined CRHoover Colorado River downstream of Hoover Dam. Producer defined CRDavis Colorado River downstream of Davis Dam. Producer defined CRParker Colorado River downstream of Parker Dam. Producer defined GunnCrystal Gunnison River downstream of Crystal Dam. Producer defined BrightAngel Bright Angel Creek in the Grand Canyon. Producer defined Latitude Decimal degrees latitude of reach location. Producer Defined 34.183444 42.026064 Longitude Decimal degrees longitude of reach location. Producer Defined -114.743441 -107.619454 ChannelUnitType Type of channel unit (i.e., microhabitat) sampled for drift invertebrates ( riffle, pool, run, cascade, side channel, or mainstem of large river). Producer Defined Mainstem Sample from the main stem channel of a river. Producer defined Run Sample from run habitat of a stream. Producer defined Date Sample date. Producer Defined Date of drift sample collection, formatted as yyyy-mm-dd. Replicate Replicate number, used to distinguish between different samples taken on the same day in the same reach of the same river. Producer Defined 1 3 MeshSize The mesh size of the drift net used to sample drift invertebrates. Producer Defined 250 250 micrometer NetWidth_Rectangular Width of the opening of the net used to sample drift invertebrates, if rectangular. Producer Defined NA No Data Producer defined 0.44 0.44 meters NetDiameter_Circular Diameter of the opening of the net used to sample drift invertebrates, if circular. Producer Defined NA No Data Producer defined 0.508 0.508 meters TimeOfDay Categorized time of day of drift invertebrate collection. Producer Defined Day Sample was collected during the day. Producer defined Dusk Sample was collected in the evening at dusk. Producer defined Dawn Sample was collected in the morning at dawn. Producer defined TimeSet Time drift net was set. Producer Defined Time, in 24-hr time, formatted as hh:mm:ss (hours:minutes:seconds) TimeCollected Time drift net was collected. Producer Defined Time, in 24-hr time, formatted as hh:mm:ss Duration Duration of drift invertebrate sampling Producer Defined 1.5 12.97 minutes DurationPrecision Whether the duration is an approximate estimation used for all samples from a project (approximate) or an exact measured time (exact) Producer Defined Exact Sample timing was collected using a watch to the nearest second at the time of sampling. Producer defined Depth Mean depth of drift of water in drift net during invertebrate sample collection. Producer Defined NA No Data. No measurement was recorded in the field at the time of sampling. Producer defined 0.1523999951232 6.40079979517441 meters Velocity Mean water velocity entering drift net during sampling. Producer Defined 0.12 2.18 meters per second VolumeSampled Total volume of water sampled for drift sample. For rectangular nets, computed as net width (from NetWidth_Rectangular column) * net height (0.25 meters) * velocity (from the Velocity column) * sample duration (from the Duration column). For circular nets, computed as net radius squared (using 0.5 * the net diameter (from NetDiameter_Circular column to determine radius) * pi * velocity (from the Velocity column) * sample duration (from the Duration column). Producer Defined 8.15 115.48 cubic meters TotalDriftAbundanceDensity Total abundance of all drift invertebrates over volume sampled. Producer Defined 0.118623962040332 170.447496677005 number of invertebrates per cubic meter of water AquaticDriftAbundanceDensity Abundance of aquatic-origin (including adults of aquatic larvae) invertebrates over volume sampled. Producer Defined 0.110782865583456 170.225963668587 number of invertebrates per cubic meter of water TerrestrialDriftAbundanceDensity Abundance of terrestrial-origin (excluding adults of aquatic larvae) invertebrates over volume sampled. Producer Defined 0.0 0.560747663551402 number of invertebrates per cubic meter of water TotalDriftBiomassDensity Total biomass of all drift invertebrates over volume sampled. Producer Defined 0.0056797583081571 11.8368098159509 milligrams dry mass of invertebrates per cubic meter of water AquaticDriftBiomassDensity Biomass of aquatic-origin (including adults of aquatic larvae) invertebrates over volume sampled. Producer Defined 0.0056797583081571 11.4932515337423 milligrams dry mass of invertebrates per cubic meter of water TerrestrialDriftBiomassDensity Biomass of terrestrial-origin (excluding adults of aquatic larvae) invertebrates over volume sampled. Producer Defined 0.0 0.78551912568306 milligrams dry mass of invertebrates per cubic meter of water NetLength Length of the net. Producer Defined 0.914 2.438 meters VelocityNet Water velocity measured within the net. Producer Defined NA No data. No measurement was recorded in the field at the time of sampling. Generally this arises from an issue with the flowmeter installed inside the net, in which case the ambient flowmeter data (VelocityAmbient column) were used for computing velocity (Velocity column). Producer defined 0.12 2.17 meters per second VelocityAmbient Water velocity measured outside and adjacent to the net. Producer Defined NA No data. No measurement was recorded in the field at the time of sampling. Generally this arises from an issue with the ambient flowmeter installed outside the net. Producer defined 0.06 2.25 meters per second Environmental Data Initiative mailing address

University of Wisconsin

Madison WI 53706 United States NA info@edirepository.org Unless otherwise stated, all data, metadata and related materials are considered to satisfy the quality standards relative to the purpose for which the data were collected. Although these data and associated metadata have been reviewed for accuracy and completeness and approved for release by the U.S. Geological Survey (USGS), no warranty expressed or implied is made regarding the display or utility of the data for other purposes, nor on all computer systems, nor shall the act of distribution constitute any such warranty. Digital Data https://doi.org/10.6073/pasta/234b23f08b48458e7d3910045ebacd9f None 20250630 Jeffrey D. Muehlbauer USGS Alaska Cooperative Fish and Wildlife Research Unit Unit Leader mailing and physical

2240 N Koyukuk Dr

Fairbanks Alaska 99709 United States 907-474-5773 jmuehlbauer@usgs.gov FGDC Biological Data Profile of the Content Standard for Digital Geospatial Metadata FGDC-STD-001.1-1999